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Matthew R. Robinson, Jill G. Pilkington, Tim Clutton-brock, Josephine M. Pemberton, Loeske E. B. Kruuk (2008). Environmental Heterogeneity Generates Fluctuating Selection on a Secondary Sexual Trait. Current Biology, 18(10), pp. 751-757, DOI: 10.1016/j.cub.2008.04.059.
